

People in the lab – Scholars, Students, Associates

Saskya
van Nouhuys
I am a population and community ecologist that works mostly on plants, their herbivores, and their parasitoids. Before coming to Indian Institute of Science (IISc), Bangalore in late 2021, I worked for 20 years in the Department of Biological Sciences at the University of Helsinki and in the Departments of Entomology and Ecology & Evolutionary Biology at Cornell University.
Scholars, Students, Associates
Postdocs & research visitors

DR. PRABITHA MOHAN (she/her)
Indian Institute of Science (2023-)
I have always been fascinated by insects and their interesting behaviours. I completed my PhD from the Central University of Kerala on the topic ‘host manipulation behaviour of parasitic wasps (parasitoids).’ After completing my PhD, I worked as a postdoctoral fellow in Long Term Ecological Observatories (LTEO) project on Arthropods at the Zoological Survey of India. There we studied the effect of climate change on Arthropods in the eco-sensitive Zones of India. As a National Postdoctoral Fellow at CES, IISc, I am working on the effect of competition on dietary specialisation in parasitoids. Besides research, I love comics, food and spending time with family and friends.

DR. SUVARNA KHADAKKAR (she/her)
Postdoc Indian institute of science (2023 & 2024)
I am interested in scarab beetle taxonomy and forest insect interactions, specifically, how forest structure shapes insect communities. For my PhD, I worked on scarab beetles around Bor Wildlife Sanctuary in Central India. I studied scarab beetle diversity, adult and larval taxonomy, seasonal distribution, variation in structure of male genitalia, developmental stages and biology of selected scarabs. I am also interested in understanding the impact of forest land diversion for non-forestry purposes on forest cover. Under NPDF I am working on Sacred Groves with dung beetles as an indicator group. Apart from this, I pursue music, Hindustani classical vocal and I am a big fan of films from all over the globe.

DR. KRISTINA KARLSSON-GREEN
Postdoc University of Helsinki (2012- 14)
Multitrophic interactions between a plant, a phytopathogen, an herbivore and a parasitoid Co-supervised with A-L Laine and Marjo Saastamoinen, University of Helsinki.

DR. CAROLINA REIGADA
Postdoctoral visitor to Metapopulation Research Group (2008)
Postdoctoral visitor to Metapopulation Research Group (2008)
Density dependent foraging behavior of a hyperparasitoid wasp. University of Helsinki. Now a professor at Depto. de Ecologia e Biologia Evolutiva – UFSCar – São Carlos, SP
Google scholar page

DR. MANASA KULKARNI (she/her)
Indian Institute of Science and Ben-Gurion University (2025-)
I have always been fascinated by cooperation in the living world. My broad research interest lies is in understanding species interactions and I have mainly worked on plant-pollinator interactions, with a focus on addressing how the partners navigate intricacies of the interaction. I completed my PhD at CES, where I worked with the fascinating system of fig trees and their pollinator wasps. I mainly studied regulation of resource flow into fig syconium (fig fruit), a space shared by developing pollinator wasp offspring and seeds of the host fig tree. Currently, I am continuing my research on oviposition behaviour in the pollinator fig wasps in a collaborative project between Ben Gurion University, Israel, and CES, IISc, under the co-supervision of Prof. Saskya and Prof. Michal. In this work, I examine the direct effects of pollination and pollinator numbers on reproductive success of pollinators.
Beside research, I am a sci-fi geek, who loves to read classics from the golden age of science fiction. Occasionally, I try my hand at writing my own sci-fi or sketching.

DR. ABHILASH NAIR
Postdoc University of Helsinki (2013- 2017)
Spatial population genetics of parasitoid wasps, University of Helsinki.

DR. DELIA PINTO ZEVALLOS
Postdoc University of Helsinki (2012)
Emissions of herbivore-induced volatile organic compounds in Plantago lanceolata and Veronica spicata. University of Helsinki. Co-supervised with J. K. Holopainen.
Currently she is a senior researcher in the Department of Environmental and Biological Sciences, Faculty of Science, Forestry and Technology, Kuopio, FI
Google Scholar page

DR. MARCELA CASTELO
Postdoc University of Helsinki (2004)
Postdoc University of Helsinki (2004)
Olfactory attraction of parasitoids to their host and host-food plant complex, University of Helsinki.
Now professor and head of the Laboratorio de Entomología Experimental – Grupo de Investigación en Ecofisiología de Parasitoides y otros Insectos (GIEP), Argentina
Google Scholar page

PROF. DIANA PÉREZ-STAPLES
Visiting professor at the Indian IISc (year 2023-2024).
I am a Professor at the Applied Ecology and Biotechnology Institute (INBIOTECA) of the Universidad Veracruzana in Mexico. I am interested in the reproduction of insect pests and vectors, the male ejaculate and female postcopulatory behaviour. My research also focuses on environmentally friendly means of pest control. At CES I am developing a protocol for the correct determination of female mating status in certain Asian fruit flies that are agricultural pests.

DR. JOANNEKE REUDLER TALSMA
Postdoc University of Helsinki (2014- 2017)
Multitrophic interactions through volatile organic chemicals. University of Helsinki Co-hosted with J. K. Holopainen

DR. NORIO HIRAI
Research visitor to the University of Helsinki (2010)
Research visitor to the University of Helsinki (2010) from Osaka University, Japan, Professor in the Graduate School of Agriculture Department of Environmental Sciences and Technology.
In our lab he worked on intraspecific competition between pupal parasitoids, University of Helsinki
Current PhD Scholars

Katta Abhishek Goud (he/him)
Indian Institute of Science (2025-)
I graduated with a BSc in Biotechnology, Zoology and Chemistry from the Osmania University and an MSc in Biochemistry and Molecular Biology from the University of Hyderabad. I have an insatiable curiosity for evolutionary biology and ecology, which was ignited during my BSc after reading the book ‘Survival Strategies’.
For my PhD, I am studying the biotic and abiotic factors that affect the symbiotic association between plants and arbuscular mycorrhizal fungi (AMF), wherein plants provide carbon to fungi in exchange for phosphorus, nitrogen and water. I am particularly interested in investigating whether different consumer groups, by exerting varying levels of herbivory, alter the amount of carbon plants allocate to fungi and thus the strength of plant-AMF associations. I will also compare these associations between agriculturally managed and unmanaged sites to test whether and how management practices like addition of nutrients and water and soil tillage affect these interactions.
Email: abhishekgoud@iisc.ac.in

Shreya Gangwal (she/her)
Indian Institute of Science (2023-)
I am interested in studying species interactions and how these interactions are shaped over time and space. I will be working on host-parasitoid interactions in butterflies and their population dynamics for my PhD. During my MS thesis, I studied how adaptation to larval crowding affected locomotory behaviour in D. melanogaster adults. I am excited about working with Prof. Saskya and learning new things in the Population and Community Ecology Lab. Besides research, I am interested in reading, dancing and bird watching.
Email: gshreya@iisc.ac.in

Ananya Kalathera (she/her)
Indian Institute of Science (2025-)
Back in 5th grade, when the teacher asked what we wanted to be when we grew up, I randomly said, “I want to be a scientist and fly around the scientific world like a butterfly.” At that time, it was mostly for attention, but years later, I ended up in a place where the research is all about plants and butterfly interactions. With a background in plant science from my bachelor’s and master’s, I have grown curious about how plants sense attacks in nature and respond to them – since unlike animals, they can’t just run away. Hoping the next few years help me explore that even more. Outside research, I had say I’m a pretty good cook and a movie lover.
Email: kananya@iisc.ac.in

Anaswar P (he/him)
Indian Institute of Science (2023-)
For me there is nothing more captivating than insects. I have always been curious about their ecology and behavior. My passion for insects has evolved from a childhood hobby of collecting them to an ambition of becoming an entomologist. Choosing Zoology for higher education helped me learn more about ecology and animal behaviour. Currently I have joined Prof. Saskya’s lab to delve into the complex world of Parasitoids and their interactions with host insects. My focus lies in the insect community ecology associated with Rutaceae plants, examining the impact of human activities on their community structure and dynamics. Beyond my scientific pursuits, I express my creativity as a shutterbug, an artist and indulge my passion for cinema as a movie enthusiast.
Email: anaswarp@iisc.ac.in

Aathira Pratap (she/her)
Indian Institute of Science (2025-)
Science has always intrigued me, understanding and deciphering the mysteries of nature is my passion. I have done my bachelor’s in zoology and microbiology and completed my masters in microbiology. I am interested in finding out associations of insects with microbes and how these associations benefit them.
I am a student of life and trying to refine myself. I love to network with people and communicate about the wondrous world of science!
Email: aathirap@iisc.ac.in

Sonali (she/her)
Indian Institute of Science (2022-)
Coming from a small town, Deoria, to this premier institute, the journey is full of excitement and uncertainties. I completed my Bachelor’s from BRD PG College Deoria where I developed an interest in plants. With the thought of exploring more in this field, I decided to pursue a Master’s in Botany from Banaras Hindu University. There I realized that I would like to continue exploring ecology more than molecular biology. And hence, I landed in the Ecology department. My research focuses on the Population and Community ecology (PoCoEco) of plants under the influence of grazers in an arid region of Karnataka dominated by tropical savannah vegetation. I am estimating the change in plant species composition and plant traits in grazed and long-term ungrazed sites in standing vegetation and soil seed bank. It’s exciting to notice the contrasting behaviour of the same species in two different conditions. Apart from research, I also love to chit-chat with people (which, most of the time, I will be seen doing) and listen to old Bollywood songs.
Email: sonali2022@iisc.ac.in
Past PhD scholars

Dr. CAMILA BERALDO
PhD, University of Helsinki (2020-2025)
“Communities ecological interactions and environmental drivers of microbiota in insect communities” Check out her dissertation here. Camila is now a postdoc at the University of Kansas

DR. JOANNEKE REUDLER TALSM
Phd, Nertherlands Institute of Ecology (2003-2007)
Phd, Nertherlands Institute of Ecology (2003-2007) (student of A. Biere & J. Harvey).
Costs and benefits of iridoids glycosides in multitrophic systems

DR. CHRISTELLE COUCHOUX
PhD, University of Helsinki (2009-13)
Intraspecific competition in the foraging behaviour of the parasitoid Hyposoter horticola PhD thesis, University of Helsinki

DR. KATIE MONTOVAN
PhD, Cornell University, Center for applied Math (2008-13)
Dissertation is titled Cooperative behavior in bees, wasps, and burglars. Now a faculty member in Math at Bennington College, USA

DR. MAARIA KANKARE
University of Helsinki (1999-2004) (primary advisor I. Hanski)
Phylogeny and host associations of Cotesia parasitoids attacking checkerspot butterflies. University of Helsinki. Maaria is a PI at Jyväskylä University
Master Students

Prankush Bansal (he/him)
Indian Institute of Science, MSc (2026-27)
Text to come soon

Pranoy MS
Indian Institute of Science (2022-)
Uses poppy plants as a model system to understand the impacts of domestication on plant morphology and defensive chemistry. He is an aspiring entrepreneur focused on innovative solutions to human problems.

Iffah Wajih
Indian Institute of Science, M.Sc. Life Sciences (2026 - )
Text and photo on their way

Chinmay Bhoyar (he/him)
Indian Institute of Science, M.Sc. Life Sciences (2025- )
Completed a Bachelor's in Zoology from Fergusson College, Pune in 2024. Curious about butterfly and caterpillar life history, especially the Red Pierrot Butterfly on host plant Kalanchoe pinnata. Has field experience with the Nature Conservation Foundation on the Satara Gecko project and with IISc tracking urban butterfly nectaring behavior in Bengaluru.
Past Master Students

Aniket Ray (he/him)
Indian Institute of Sciences, MSc (2025-26
His MS thesis is titled "Dynamic economic threshold modeling of stage-structured pest–natural enemy dynamics," which incorporates age structure and a type II Hollings response into a baseline dynamic sprey threshold model.
Email: aniketray@iisc.ac.in

Pheba Anna Biju (she/her)
St. Josephs University, MSc project at IISc (Dec. 2025 - April 2026)
Conducted a comparative assessment of Plant Lepidoptera Parasitoid communities on the Rutaceae family across varying land management intensities (co-advised with Anaswar P).

Narshiman N. (he/his)
Indian Institute of Sciences, MSc Life Sciences (Aug. 2023 - May 2025)
His MSc thesis was "Reproductive strategies in Asteraceae in the context of invasion ecology and evolution." Moving on to continue working on plant reproductive ecology with Adrienne Nicotra at Australian National University.

Yann Bailly
MSc intern from Lyon, France (2017)
Investigated sex determination in the parasitoid Cotesia glomerata and the costs of parasitism for developing caterpillars at Helsinki.

Aapo Ahola
University of Helsinki, MSc (2011 - 13)
Researched the heritability of traits associated with landscape structure in Plantago lanceolata (co-advised with Anna-Liisa Laine).

Dr. David Muru
MSc project from France (2012)
Investigated the role of group behavior on the relationship of the butterfly Melitaea cinxia with the parasitoid H. horticola at Helsinki.

Markku Ojanen
University of Helsinki, MSc (05 - 08)
Explored plant, phytopathogen, and parasitoid interactions (co-advised with Anna-Liisa Laine).

Dr. Riikka Kaartinen
University of Helsinki, MSc (03 - 06)
Investigated spatial foraging behavior of the parasitoid Hyposoter horticola, publishing a paper in Proceedings of the Royal Society B (2008). She currently works as a journalist for Suomen Luonto.

Dr. Christian Anton
MSc project Jena, Germany (00 - 01)
Studied behavioral and developmental responses of a specialist parasitoid wasp to host larvae feeding on two host food plant species at Helsinki. Currently works for the National Academy of Sciences Leopoldina in Germany.

Thejus Manohar (he/him)
JSS Academy of Higher Education and Research, MSc project at IISc (2026)
His Thesis project is titled "Taxonomic identification of Parasitoid wasps of Lepidoptera on Rutaceae in Southern India Using Molecular approaches"
He worked on molecular identifcation and taxonomic classiciation of some of the many egg parasitoids of Lepidotera collected as part of our study of the Lepidoptera food web associated with wild and domesticaated Rutaceae,

Vandana P M (she/her)
Yuvaraja's College, University of Mysore, MSc project at IISc 2025
Studied sources of mortality of Lepidoptera eggs on sentinel egg cards on the IISc campus (co-advised with Prabitha Mohan).

Rajnish Shree Raj
IISER Tirupati, MSc project IISc (June 2024 - May 2025)
His MSc project was on "Foraging behavior of Trichogramma chilonis on Corcyra cephalonica egg under competitive and Non-competitive conditions." Currently continuing deeper work into wasp behavior (co-advised with Prabitha Mohan).

Dr. Pil Rasmussen
University of Helsinki, MSc (2013 - 14)
Researched multitrophic interactions between a plant, herbivore, and parasitoid, resulting in a publication in Ecological Entomology (Rasmussen et al 2017). Currently works at the National Gallery of Denmark.

Terhi Lahtinen
University of Helsinki, MSc student (2010 - 2012)
Studied the variation in the response to drought in Plantago lanceolata.

Riikka Alanen (Hartikainen)
University of Helsinki, MSc (2010 - 2011)
Researched habitat fragmentation and the germination of P. lanceolata and V. spicata (co-advised with Anna-Liisa Laine).

Anne Reichgelt
University of Helsinki, MSc (07)
Researched density dependent parasitism by the hyperparasitoid Mesochorus stigmaticus.

Dr. Lucie Salvaudon
MSc project Paris, France (2002)
Conducted an experimental test of host avoidance of parasitism by Hyposoter horticola at Helsinki. Currently works at the Pierre and Marie Curie University in Paris.

Eeva Punju
University of Helsinki, MSc (00 - 02)
Investigated interspecific competition between immature parasitoid wasps within host larvae, leading to a publication in Oikos (2010).

Gauravya Mohan (any pronouns)
Indian Institute of Science, MSc Life Sciences 2024- 2026
Gauravya's MSc thesis: Exploring the Interaction of Ants with their Resident Trees in an Indian Savannah
email:gauravyamohan@gmail.com
He worked out the network of ants associated with homotera on different tree species at our Challakere field site.

Shri Avanthika (she/her)
Central University of Karnataka, M.Sc. project IISc 2025
Researched moth communities associated with wild and cultivated Rutaceae (co-advised with Anaswar P).

Dr. Emma Hakanen
University of Helsinki, MSc (2014 - 2017)
Studied the mating structure of the parasitoid Cotesia melitaearum (co-advised with A. Nair).

Marina Granger-Brown
Harper Adams University, UK, MSc (2014)
Studied intraspecific competition by the parasitoid wasp Hyposoter horticola at Helsinki.

Peter Kullberg
University of Helsinki, MSc (2010 - 12)
Studied habitat fragmentation and the performance of P. lanceolata and V. spicata (co-advised with Anna-Liisa Laine).

Suvi Niemikapee (Pakarinen)
University of Helsinki, MSc (2010 - 2011)
Studied host-parasitoid relationships in different Cotesia melitaearum and Melitaea cinxia populations around the Baltic Sea, resulting in a publication in Insects (2012).

Kaisa Torri
University of Helsinki, MSc (2005 - 07)
Studied the natural history of the butterfly Melitaea athalia and its parasitoids in Åland, Finland, resulting in a publication in Arthropod-Plant Interactions (2008).

Johanna Ehrnsten
University of Helsinki, MSc (01 - 03)
Studied the movement and oviposition behavior of the mobile parasitoid wasp Hyposoter horticola, publishing in Behavioral Ecology (2004).
Interns & Undergraduates

SHUBHANGI SHANDILYA INTERN (she/her)
Central University (BBAU) Lucknow�
I am working on a butterfly diversity project, where I analyze year-long data collected from three sites. The dataset includes records of butterfly species observed along with environmental parameters such as temperature, humidity, rainfall, and wind speed. Using R for data analysis, I study how these factors influence butterfly diversity and abundance.
June 2025 -?

SURAJ KUMAR INTERN (he/him)
Vinoba Bhave University
Jan. 2026 - May 2026

PRANKUSH BANSAL UG INTERN (he/him)
Indian Institute of Science
Biology major pursuing a BSc. Interested in Behavioural and community Ecology. Worked on Dung Beetle Morphology and Diversity with Dr. Suvarna Khadakkar. Started a UG project in Kavita Isvarans lab on life history trait variation.
2024 & 2025

ANIKET RAY - UG THESIS STUDENT (he/him)
Indian Institute of Science
Did his undergraduate project on modeling aspects of species invasion, titled Exploring Role of Propagules in Invasion Biology through Literature Review and Simple Simulations. Now working on his MSc modeling insecticide effects.
May 2023 - 2025

NITHINSAKTHI T - UG Thesis student
Indian Institute of Science
Undergrad senior thesis project involving a literature survey of modeling foraging behavior of parasitoids under competition and where resources are ephemeral.
Jan - May 2023

NIKHIL KOLLINS INTERN (he/him)
IISER TVM
BS-MS graduate broadly interested in entomology and evolutionary ecology. Currently working with PhD student Sonali on Grassland ecology and insect identification from IISc Challakere campus.
July 2025 - May 2026

NANDURAJ S INTERN
IISER TVM
Completed graduation and is interested in studying plant-pollinator interactions. Currently working with PhD student Anaswar on the oviposition preferences of butterflies and the performance of butterfly caterpillars on various host plants.
July 2025 - Jan 2026

Snata Chakraborty PhD visitor (she/her)
University of Portsmouth, UK
Collaborating to learn life history of parasitoids. Studying host-parasitoid interactions between an Eurytomid wasp and the tropical tent web spider. Holds an Anthropology background (U. of Calcutta, U. of Delhi) and an MSc in Evolution, Ecology, and Systematics from LMU Munich.
Aug - Sept 2025

PRASHANT DALAWAI - THESIS STUDENT
College of Forestry, Ponnampet, Karnataka
Final year BSc student working as an intern on the project by Dr. Suvarna Khadakkar. Focuses on Dung beetle diversity in Sacred Groves and Coffee plantations in Kodagu district of Karnataka.
2023 - 2024

ADITYA NAYAK - Summer Intern
IISER Mohali
BS-MS student who worked on finding parasitoids of Lepidoptera on Citrus, and conducted a literature review of change in genome size associated with plant domestication.
2022

BIPASHA DAS INTERN (she/her)
IISER Kolkata
BS-MS graduate currently working on the plant feeding behaviour of mosquitoes under Dr. Prabitha Mohan. Previously worked on host-parasitoid interactions. Interested in insect chemical and behavioural ecology and ecophysiology.
Jan. 2026 - May 2026

Monika Gandhi INTERN
Postgraduate in Zoology interested in insect ecology, taxonomy, life history observations, and plant-pollinator interactions. Worked with PhD student Sonali on insect identification from grasslands of IISc Challakere campus.
Sept. 2025 - Dec. 2025

Anton Toms MSc Intern
Christ University, Bangalore
Worked on the behavior of foraging Trichogramma chilonis with MSc project student Rajnish, and on the fate of Corcyra cephalonica eggs in the field with MSc project student Vandana.
Dec 2024 - June 2025

CHETHANA NAIR - UG SUMMER INTERN
Indian Institute of Sciences
Second year-completed undergrad exploring ecology. Interests align with the lab in population ecology as well as community-wide interactions.
2023
Project Associates and Assistants

PRAKRUTI MEHTA
Project associate (she/her) (Feb 2024-)
With a fascination in ecology, and interest in the environment, I completed my master’s in environmental science. With a diverse background in studying birds, butterflies, macro fungi and invasive snails, I have honed my skills in ecology and conservation. Currently, at the PoCo Eco lab, I am delving deeper into the community ecology of parasitoids, helping with sample collection, and rearing insects, including butterflies and parasitoids. I am eager to understand the complexities of these ecosystems and the effect of biotic and abiotic factors on these communities. In addition to my scientific curiosity, I’m a cat parent to two adorable fur babies and a major bookworm.
Indian Institute of Science

Malay Pandey
Project Associate, (April-Oct 2025)
During his B.Sc. at Azim Premji Uni., in Bangalore, Malay explored habitat selection by cockroaches. For his M.Sc. thesis, in Ben-Gurion Uni., he explored foraging responses and physiological consequences of crude-oil pollution on gerbils. Additionally, he worked on the social dynamics of Free-Ranging Dogs in Bengaluru. He also studied maternal investments in egg sac construction by widow spiders. In the PoCoEco Lab, he worked on the Asian Citrus Psyllid project studying monitoring methods, and the natural enemy community.
As of Nov., 2025 he is a PhD student at the University of Turin in Italy studying cricket behavior.
Indian Institute of Science

Girish G.
(Sept. 2024-), Assistant
Girish wears many hats in the lab, from office work to transportation, to help with research. One day he may be going around campus chasing down signatures and the next day he’ll be helping a student collect black buck droppings from the field for her research.
Indian Institute of Science

Vanshika Pal
Research assistant, (May 2023 – July 2024)
Vanshika Pal completed her education till post grad from University of Lucknow. I In PoCo Eco she surveyed butterfly behaviour and diversity as a function of weather, and worked on behavior of Trichogramma parasitoid wasps with respect to competition. Now she is a PhD scholar at Ahmedabad University.
Indian Institute of Science

Tapasya Thapa
Project Associate, (March 2022- October 2025)
Tapasya was brought up in a small city, Dehradun. She did her MSc in Biotechnology in 2018 and joined the Wildlife Institute of India, Dehradun where she gained experience in conservation genetics tools and techniques. She started working in the PoCoEco lab when it was just an empty room in 2022. Since then she has had many roles in the lab, including the molecular phylogenetic work toward identifying the diverse parasitoids associated with Lepidoptera feeding on Ructaceae.
As of Nov., 2025 she is PhD student at the University of Tartu in Estonia.
Indian Institute of Science